Phylogenetic position of the crocodylian Megadontosuchus arduini (De Zigno, 1880) and tomistomine palaeobiogeography

A cladistic analysis of Megadontosuchus arduini from the middle Eocene of Monte Duello (NE Italy) confirms its tomistomine relationships, but the low number of scorable characters determines a low resolution within the tomistomine clade. However, Megadontosuchus is clearly distinct from the other Eocene European or North African tomistomines, in having a moderate elongated but robust rostrum, massive maxillary and dentary teeth and large supratemporal fenestrae. The rostrum and teeth characteristics could indicate that M. arduini had a degree of feeding specialization intermediate between Maroccosuchus zennaroi and the Eocene European tomistomines. A summary of tomistomine palaeobiogeo− graphy suggests that despite only one species with a rather restricted range survives at present, such a clade had a glorious past with a world wide distribution documented by a conspicuous fossil record that starts at least in the early Eocene. At present, a detailed knowledge of tomistomine palaeobiogeography is hindered by the lack of modern taxonomic revisions of some fossil remains and therefore by the poor understanding of phylogenetic relationships.